Muthukumarasamy Arunachalam、Manickam Raja、Subramaniyam Nandagopal、Appukuttan Chandran
????? Puntius chalakkudiensis ??????? Puntius denisonii (Day 1865) ???1999 ? Menon?Rema Devi ? Thobias ?????????? Chalakkudy ??????????????????? Achankoil ?????? [1] ????????????????????P. chalakkudiensis ? P. denisonii ?????????????????????P. denisonii ???????????? Pathanamthitta ? Plappally ? Pamba ???? Puntius chalakkudiensis ???????????
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